Vertical gap appearing in the second column of a multicolumn environment (oxford bioinformatics template)

Question

I have a problem with a vertical gap appearing in the second column of the second page, instead of text/figure. This problem is not exclusive to the pages near the bibliography because this gap also appeared in another page. Please advise on how I can pull up the text/figure to the top of second column (I'm using the oxford bioinformatics template https://www.overleaf.com/latex/templates/template-for-oxford-bioinformatics-journal-new-version/zjrmbrmtrytg).
PassOptionsToPackage{utf8}{inputenc}
documentclass{bioinfo}
copyrightyear{2015} pubyear{2015}
usepackage{comment}
usepackage[caption=false]{subfig}
usepackage{lipsum}

access{Advance Access Publication Date: Day Month Year}
appnotes{Manuscript Category}

begin{document}
%firstpage{1}

lipsum[90-100]

begin{figure}[!tpb]%figure1
fboxsep=0ptcolorbox{gray}{begin{minipage}[t]{235pt} vbox to 100pt{vfillhbox to
235pt{hfillfontsize{24pt}{24pt}selectfont FPOhfill}vfill}
end{minipage}}
%centerline{includegraphics{fig01.eps}}
caption{Caption, caption.}label{fig:01}
end{figure}

begin{figure}[!tpb]%figure1
fboxsep=0ptcolorbox{gray}{begin{minipage}[t]{235pt} vbox to 100pt{vfillhbox to
235pt{hfillfontsize{24pt}{24pt}selectfont FPOhfill}vfill}
end{minipage}}
caption{Caption, caption.}label{fig:01}
end{figure}

begin{figure}[!tpb]%figure1
fboxsep=0ptcolorbox{gray}{begin{minipage}[t]{235pt} vbox to 100pt{vfillhbox to
235pt{hfillfontsize{24pt}{24pt}selectfont FPOhfill}vfill}
end{minipage}}
caption{Caption, caption.}label{fig:02}
end{figure}

begin{figure}[!tpb]%figure1
fboxsep=0ptcolorbox{gray}{begin{minipage}[t]{235pt} vbox to 100pt{vfillhbox to
235pt{hfillfontsize{24pt}{24pt}selectfont FPOhfill}vfill}
end{minipage}}
caption{Caption, caption.}label{fig:03}
end{figure}

section*{Acknowledgements}

Text Text Text Text Text Text  Text Text.  citealp{Boffelli03} might want to know about  text
text text textvspace*{-12pt}

section*{Funding}

This work has been supported by the... Text Text  Text Text.vspace*{-12pt}

bibliographystyle{natbib}

%bibliography{document}

begin{thebibliography}{}

bibitem[Bofelli {it et~al}., 2000]{Boffelli03}
Bofelli,F., Name2, Name3 (2003) Article title, {it Journal Name}, {bf 199}, 133-154.

bibitem[Bag {it et~al}., 2001]{Bag01}
Bag,M., Name2, Name3 (2001) Article title, {it Journal Name}, {bf 99}, 33-54.

bibitem[Yoo textit{et~al}., 2003]{Yoo03}
Yoo,M.S. textit{et~al}. (2003) Oxidative stress regulated genes in nigral dopaminergic neurnol cell: correlation with the known pathology in Parkinson's disease. textit{Brain Res. Mol. Brain Res.}, textbf{110}(Suppl. 1), 76--84.

bibitem[Lehmann, 1986]{Leh86}
Lehmann,E.L. (1986) Chapter title. textit{Book Title}. Vol.~1, 2nd edn. Springer-Verlag, New York.

bibitem[Crenshaw and Jones, 2003]{Cre03}
Crenshaw, B.,III, and Jones, W.B.,Jr (2003) The future of clinical cancer management: one tumor, one chip. textit{Bioinformatics}, doi:10.1093/bioinformatics/btn000.

bibitem[Auhtor textit{et~al}. (2000)]{Aut00}
Auhtor,A.B. textit{et~al}. (2000) Chapter title. In Smith, A.C. (ed.), textit{Book Title}, 2nd edn. Publisher, Location, Vol. 1, pp.???--???.

bibitem[Bardet, 1920]{Bar20}
Bardet, G. (1920) Sur un syndrome d'obesite infantile avec polydactylie et retinite pigmentaire (contribution a l'etude des formes cliniques de l'obesite hypophysaire). PhD Thesis, name of institution, Paris, France.